Why does port forwarding and uPnP increase P2P download speeds?

LuckyTommy

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So I know what they are: port forwarding lets computers on the internet connect directly to the PC behind my router; and turning on UPnP lets devices automatically connect to my network ( hence plug and play ).

P2P works just fine but slowly when these settings are off. Then I enable these settings, speeds increase dramatically, can anyone explain why?
 
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