Well, you say it has spark and you gave it fuel. This leaves breathing (intake/exhaust) and timing. You are testing the spark in open air, what tool are you using? Can you vary the gap?
The combustion chamber provides a great deal of resistance to the spark, it may take an easier path when it is presented with compression pressures.
I would expect your timing is thru gears, you would also hear differences in cranking since compression would not be up to par.
Look more closely at your spark.